Amphibians are a group of cold-blooded vertebrates that include frogs, toads, salamanders, and caecilians. They are characterized by their ability to live both in water and on land, a trait that is reflected in their name: "amphibian" means "double life." Amphibians first appeared during the Devonian period, around 370 million years ago, making them one of the oldest groups of land vertebrates. They evolved from lobe-finned fish, which possessed fleshy, lobed fins with bones similar to those in the limbs of modern amphibians.
  • Amphibians include frogs, toads, salamanders, and caecilians.
  • They are vertebrates adapted to both aquatic and terrestrial environments.
  • The first amphibians appeared about 370 million years ago.
  • Their ancestors were lobe-finned fish, which had limb-like fins.
Amphibians first appeared during the Devonian period, around 370 million years ago.
Amphibians evolved from lobe-finned fish.

Transition to Land

The evolutionary transition from water to land required significant adaptations. Early amphibians developed lungs for breathing air, as well as limbs capable of supporting weight on land. Their skin became more resistant to desiccation (drying out), although it remained permeable to water and gases, making them vulnerable to environmental changes. Amphibians also retained a connection to water for reproduction, as their eggs lack a protective shell and must be laid in moist environments.
  • Early amphibians evolved lungs for breathing air.
  • Limbs replaced fins to support movement on land.
  • Skin became more resistant to drying but remained permeable.
  • Reproduction still depended on water due to egg characteristics.
Lungs and supportive limbs were crucial for the transition to land.
Amphibian eggs lack a protective shell, so they must be laid in moist environments to prevent drying out.

Major Fossil Groups

Ichthyostegalians are among the earliest known tetrapods and are considered close relatives of modern amphibians.
Ichthyostegalians are among the earliest known tetrapods and are considered close relatives of modern amphibians. They had both fish-like and amphibian-like features, such as jaws and gills (fish-like) along with limbs and lungs (tetrapod traits). Fossils of Ichthyostegalians, dating from about 365 million years ago, show transitional characteristics that bridge the gap between fish and amphibians.
Proposed ancestral group including modern amphibians and some extinct relatives: Lissamphibia (modern amphibians), and extinct groups like Temnospondyli and Lepospondyli.
GroupDescriptionTime PeriodSignificance
IchthyostegaliansEarly tetrapods with mixed featuresLate DevonianTransitional fossils
TemnospondyliDiverse, some giant amphibiansPaleozoic–MesozoicPossible ancestors of lissamphibians
LepospondyliSmall, varied formsCarboniferous–PermianPossible relatives, some extinct
Ichthyostegalians were early tetrapods showing both fish-like and amphibian-like characteristics.

Modern Amphibians (Lissamphibia)

Modern amphibians belong to the subclass Lissamphibia, which includes all living species of frogs (Anura), salamanders (Caudata or Urodela), and caecilians (Gymnophiona). This group first appeared in the fossil record during the Triassic period (about 250 million years ago) and is characterized by smooth, moist skin, specialized ear structures, and distinct larval stages.
  • Lissamphibia includes all living amphibians.
  • Major groups: Anura (frogs/toads), Caudata (salamanders), Gymnophiona (caecilians).
  • First appeared in the fossil record during the Triassic.
  • Characterized by smooth skin and complex life cycles.
Lissamphibia includes frogs, salamanders, and caecilians.
Modern amphibians appeared in the Triassic period.

Conclusion

Amphibians are an ancient group of vertebrates that first emerged during the Devonian period, evolving from lobe-finned fish. Their unique adaptations for life both in water and on land—such as lungs, limbs, and specialized skin—enabled them to occupy new ecological niches. Modern amphibians (Lissamphibia) include frogs, salamanders, and caecilians, and they trace their origins to the Triassic period.
  • Amphibians evolved from lobe-finned fish about 370 million years ago.
  • Key adaptations for land included lungs, limbs, and resistant skin.
  • Modern amphibians belong to Lissamphibia and diversified in the Triassic.